E. L. Wehner-Welch Foundation Chair in Chemistry. Regents Professor. Distinguished Professor of Chemistry, Distinguished Professor of Molecular and Cellular Medicine, and Professor of Biochemistry and Biophysics. B. S., 1964, California Institute of Technology. Ph. D., 1973, University of Oregon. Helen Hay Whitney Fellow, 1974-1976, Columbia University. Biochemistry and Biophysical Chemistry. Molecular mechanisms involved in protein translocation across and integration into various membranes and in cell lysis by bacterial toxins; protein-membrane, protein-protein, and protein-RNA interactions; fluorescence spectroscopy and resonance energy transfer; photocrosslinking.